Mawle was born in Oxford. He grew up in a Victorian manor house on a farm outside Shipston-on-Stour, Warwickshire. He is one of three children from a family with a history of nine generations of farming in Warwickshire. His mother was a teacher. His father Richard still farms the land. He attended The Croft prep schoolon the outskirts of Stratford-upon-Avonwhere he was diagnosed as severely dyslexic and as a result he went from the ages of 13 to 16 at a boarding school for special needs. Mawle left the school at 16 to become an actor, but contracted labyrinthitis, leaving him 70 per cent hearing-impaired in the upper register and with tinnitus. He persuaded the director of the local college at Stratford-upon-Avon to let him study for a BTEC in performing arts while doing an assortment of jobs such as fitness instructor and dish-washer. He also did some acting for the Box Clever Theatre Company and in his own production ofSolo Hamlet. He won a scholarship to study at the Bristol Old Vic's theatre school. Mawle graduated from Bristol Old Vic Theatre Schoolin 2002 in the same year he appeared in the Irish HurlingTV adverts for Guinness. He left in 2002 with no agent and worked as an assistant in a special needs school. He landed his first major role in a 2003 production of Troilus and Cressidaplaying Troilus for Shakespeare at The Tobacco factory, work at the Nuffield Theatre, Southampton, in Hamlet and in 2005 the Royal Exchange Manchester's production of Antony & Cleopatrafollowed. It was his performance in the television film Soundproofin 2006 that made his biggest impact where a deaf actor was needed for the lead role. Mawle was nominated for the RTS Breakthrough on Screen Awardand director Ed Coulthard won the BAFTAfor best director in 2007. He has appeared in a number of British television productions since including Dunkirkwinner of the Huw Wheldon BAFTA Award for Factual Drama, and in Adrian Shergold's Persuasionand Channel 4's gay drama Clapham Junctionin the role of Tim. Time Outsaid "Mawle and Treadaway put in Bafta-winning efforts". In 2008 he starred as Jesus in BBC/ HBO Filmsminiseries of The Passion, and as a guest star in Foyle's War"Broken Souls" episode. In 2008 Mawle returned to the theatre in The Last Days of Judas Iscariotat the Almeida Theatredirected by Rupert Goold. Listed for Outstanding Newcomer by The Evening Standard Theatre Awardsand nominated for WhatsOnStage Theatre Choice Awardsfor Best Newcomer for his performance as Judas. In 2009 Mawle featured in films Nineteen Eighty(part of the Red Riding Trilogy) directed by James Marshand in Heartlessby Philip Ridley and in the spring in Dominic Savage's Freefallopposite Dominic Cooperand Aidan Gillen. In the same year, he starred in episode 4 of Jimmy McGovern's The Street. In 2010 Mawle featured in artist David Austen'sEnd of Loveand in the short filmSometimes The Moon Is Velvet, which toured festivals. He also appeared in Agatha Christie's Murder on the Orient Express, BBC drama Five Daughters, Dominic Savage's Diveand Made in Dagenhamstarring Sally Hawkins. He also starred with Rosamund Pikeas lovers Gerald Crich and Gudrun Brangwen in the 2011 BBC Fourtelevision adaptation of Women in Love. Mawle currently stars as Benjen Stark in the HBO series Game of Thronesadapted from George R. R. Martin's A Song of Ice and Firenovels. He appeared in several 2012 projects including the Working Title production of Sebastian Faulks's acclaimed novelBirdsong, the Bruce Willisfilm "The Cold Light of Day", the Tim Burton-produced Abraham Lincoln: Vampire Hunter, and the independent British filmShell. In 2011, he was featured in The Horrible Crowes' music video for their debut single "Behold The Hurricane".